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Lynch, J. D., & Sliárez-Mavorga, Á. M. (2001). The distributions of the gladiator frogs (Hyla Boanas group) in Colombia with coments on size variation and sympatry. Caldasia, 23(2), 491-507.

https://revistas.unal.edu.co/index.php/cal/article/download/17719/18562/56745

Five species of gladiator frogs (the Hyla boans group) occur in Colombia. Hyla wavrini barely enters Colombia (it is confined to the extreme eastern edge ofthe country) and H. rosenbergi is endemic to the biogeographic Chocó. The other three species are more widely distributed. Hyla boans is found in the Chocó, Middle Magdalena, and Amazonas-Orinoquia below 1QOO m. “Hyla crepitans” is found in the Magdalena drainage as well as the western edge ofthe Orinoquia and less commonly along the Caribbean coast and in the Maracaibo Basin. This species occurs from sea level to more than 2000 m. Hyla pugnax occurs across the Caribbean lowlands and up the Magdalena valley as well as in the Maracaibo Basin and at a few cis-Andean sites, but always below 500 m. Sympatry is relatively common between H. boans and H. rosenbergi but, less common between H. boans and either “H. crepitans” or H. pugnax. Maximal sympatry results in sympatry of H. boans, H. pugnax, and H. rosenbergi – normally no more than two species are sympatric. Sexual dimorphism in size is evident in the smallest species (“H. crepitans “) but not in the others. AH exhibit geographic variation in size, perhaps related to the combination ofsympatric species ofthe group.